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- ArticleSrivastava PN, Akruk SR, Williams WL.J Exp Zool. 1979 Mar;207(3):521-9.The acrosomal extracts from rabbit spermatozoa were more effective in dissolving the zona pellucida of the rabbit ova in the absence than in the presence of Ca2+. The extracts had high proteolytic activity but low esterolytic activity at alkaline pH in which pH effective zona hydrolysis occurred. The results suggest that acrosomal proteinases in addition to acrosin may be involved in the zona penetration.